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These crates contain all different resources you'll need to tune up your Toons: Regional Materials of all rarities and Gold. They come in different tiers and there's one per Region (except for Tasmania, as these region does not feature this type of crates).
To get a higher tier crate you need to earn trophies in brawl and get into a higher league. You can always check each crate to see the selection of rewards you could get from it.